Scleropodium occidentale B.E. Carter

Scleropodium occidentale is a medium-large, green to yellow-green, pleurocarpous moss, which forms large mats on rocks and occasionally tree bases in and directly adjacent to seasonal creeks and streams. Its irregularly spaced branches taper slightly toward the tips and appear quite swollen and poofy even when dry. Its leaves are concave and the midrib ends in a spine, a feature which helps to differentiate it from the closely related S. obtusifolium.
Scroll down for more photos (click to enlarge) and descriptions of habitat, plants, leaves, distribution map, name origin, and similar species.
Scleropodium occidentale
Scleropodium occidentale (CAH #791). CC BY-NC Amanda Heinrich
Habitat: Scleropodium occidentale is strictly an aquatic or semi-aquatic species, growing on rock or occasionally tree roots, in or directly adjacent to and thus in the splash zone of creeks or seasonal streams.
Scleropodium occidentale
Scleropodium occidentale moist on small boulder in seasonal stream, Santa Ynez Mountains, Santa Barbara County, CA (CAH #791) CC BY-NC Amanda Heinrich
Scleropodium occidentale
Scleropodium occidentale dry on boulder in seasonal stream, Santa Ynez Mountains, Santa Barbara County, CA (CAH #796) CC BY-NC Amanda Heinrich
Habit:  Scleropodium occidentale forms extensive messy mats, which obscure its irregularly spaced branching pattern. The branches taper slightly toward the tips and appear quite swollen and poofy even when dry.
Scleropodium occidentale
Scleropodium occidentale moist, Santa Ynez Mountains, Santa Barbara County, CA (CAH #791) CC BY-NC Amanda Heinrich
Scleropodium occidentale
Scleropodium occidentale dry, Santa Ynez Mountains, Santa Barbara County, CA (CAH #796) CC BY-NC Amanda Heinrich
Plants:  Scleropodium occidentale plants are medium-large (to 5+ cm), with many irregularly spaced branches which tend to bend midway and taper slightly at the tips (attenuate). The concave leaves are closely spaced on the stem, and overlap neatly when dry (julaceous), but are not exceedingly tightly appressed to the stem, resulting in a poofy appearance (tumid).
Scleropodium occidentale
Scleropodium occidentale plant moist (CAH #792) CC BY-NC Amanda Heinrich
Scleropodium occidentale
Scleropodium occidentale plant dry (CAH #792) CC BY-NC Amanda Heinrich
Leaves: Scleropodium occidentale leaves are ovate (sometimes more lanceolate) and concave, with a rather strong costa extending >70% up and terminating in a spine, a feature which helps to distinguish it from the closely related Scleropodium obtusifolium. The leaf margins are often slightly serrated and inrolled in the upper part of the leaf and the tips are generally acute.
Sporophytes:  Please see the references below for information on Scleropodium occidentale sporophytes as I have yet to encounter them in Santa Barbara County.
Distribution:  Scleropodium occidentale is found in California primarily west of the Sierra Nevada, into Oregon and BC, with a few scattered collections to the east. In Santa Barbara County thus far I have found it primarily in creeks on the north side of the Santa Ynez Mountains and have yet to find a creek containing both Scleropodium occidentale and Scleropodium obtusifolium.
Scleropodium occidentale distribution
 Scleropodium occidentale collections. Map generated by google maps using data from the Consortium of Bryophyte Herbaria of accessioned collections from herbaria.
Similar Species in Santa Barbara County
Scleropodium obtusifolium has a nearly identical habitat preference and appearance but its leaves have a shorter, weaker costa, which lacks a terminal spine.
Name: The genus name Scleropodium is derived from the Greek word skleros, meaning hard, and podion, meaning foot, and refers to the stiff seta. The species epithet, occidentale is a reference to its distribution in western N. America.
References and Further Reading
Systematics and ecology of the moss genus Scleropodium (Brachytheciaceae), Benjamin Elias Carter, PhD Dissertation
Scleropodium occidentale, Michael S. Ignatov, Flora of North America Vol. 28, www.efloras.org