Mosses, hornwort, liverwort
Click photos to enlarge. All photos CC BY-NC Amanda Heinrich.
LiverwortsMossesHornworts
Species Worldwide7,00016,000-20,000200
Species in California1306508
Species in Santa Barbara County231305
Gametophyte MorphologyComplex thallose, simple thallose, bottlepore and leafy liverwortsAcrocarpous and pleurocarpous mossHornwort
Sporophyte MorphologyLiverwort sporophytes: transient, with spherical capsules on flimsy stalk; split open along lines of dehiscence
Liverwort sporophytes
Moss sporophytes: sturdy, persistent, typically with complex opening

Moss sporophytes
Hornwort sporophytes: horn-shaped; split open like banana

Hornwort sporophytes
Costa (midrib on leaf)No costa on leaves
Liverwort leaves no costa
Costa present on leaves of most species
Moss leaf with costa
No costa
Elaters
(spring-like cells, which aid in spore dispersal)
Elaters typically present
Liverwort elaters
No elatersPseudoelaters (non-functional)
Hornwort pseudoelaters
Rhizoids
(attach plant to substratum)
Unicellular rhizoids
Liverwort rhizoids unicellular
Multicellular rhizoids
Multicellular moss rhizoids
Unicellular rhizoids
Hornwort rhizoids unicellular
ChloroplastsMany small chloroplasts per cell
Liverwort leaf cells with many small chloroplasts
Many small chloroplasts per cell
Moss leaf cells with many small chloroplasts
One or two giant chloroplasts per cell
Hornwort chloroplasts
Oil bodiesMany liverworts contain oil bodies; shape, number and size may aid in species identification
Leafy liverwort oil bodies
No oil bodiesNo oil bodies