Scleropodium obtusifolium (Mitten) Kindberg

Scleropodium obtusifolium is a medium-large, glossy light green to yellowish pleurocarpous moss, which forms thick shaggy mats and extensive carpets on rocks and soil, in and immediately adjacent to creeks and streams. Its irregularly spaced branches are disorganized and appear quite swollen and poofy even when dry. The leaves are deeply concave and rounded at the tip, and the midrib does not end in a spine, a feature which helps to differentiate it from the closely related S. occidentale.
Scroll down for more photos (click to enlarge) and descriptions of habitat, plants, leaves, sporophytes, distribution map, name origin, and similar species.
Scleropodium obtusifolium
Scleropodium obtusifolium (CAH #1115). CC BY-NC Amanda Heinrich
Habitat:  Scleropodium obtusifolium is strictly an aquatic or semi-aquatic species, growing on rock or soil or occasionally tree roots, in or directly adjacent to creeks or seasonal streams.
Scleropodium obtusifolium habitat wet
Scleropodium obtusifolium on rock in seasonal streamlet, wet (CAH #1115). CC BY-NC Amanda Heinrich
Scleropodium obtusifolium habitat dry
Scleropodium obtusifolium on thin soil over rock in seasonal streamlet, dry (CAH #1003). CC BY-NC Amanda Heinrich
Habit:  Scleropodium obtusifolium, with its plentiful but irregularly spaced and sized branches, forms messy, shaggy mats or carpets. Its tightly spaced concave leaves spread from the stem when moist, accentuating the furry look. When dry, the leaves are held more tightly to the stem but retain their concave shape, resulting in a poofy (tumid) wormlike (julaceous) appearance.
Scleropodium obtusifolium
Scleropodium obtusifolium moist (CAH #1115). CC BY-NC Amanda Heinrich
Scleropodium obtusifolium
Scleropodium obtusifolium dry (CAH #1003). CC BY-NC Amanda Heinrich
Plants:  Scleropodium obtusifolium plants are medium-sized (3-5 cm in SB County, up to 10 cm), with many irregularly spaced branches which often arch upward, and do not taper significantly at the tips (not attenuate). The concave leaves are closely spaced on the stem, overlapping neatly when dry (julaceous), but not exceedingly tightly appressed to the stem, resulting in a poofy appearance (tumid).
Scleropodium obtusifolium
Scleropodium obtusifolium moist (CAH #1115). CC BY-NC Amanda Heinrich
Scleropodium obtusifolium with sporophytes
Scleropodium obtusifolium dry with sporophytes (CAH #1003). CC BY-NC Amanda Heinrich
Leaves: Scleropodium obtusifolium leaves are deeply concave, typically ovate, with rounded or obtuse tips (sometimes more acute). The margins are mostly smooth but sometimes minutely toothed (serrulate) near the tips, and often inrolled. The midrib (costa) tapers gradually and generally peters out 70-85% of the way up the leaf. Unlike most Scleropodium species, the costa does not end in a spine, and this feature helps to distinguish it from the closely related Scleropodium occidentale.
Sporophytes: Scleropodium obtusifolium sporophytes have capsules which are generally held at a horizontal angle or dangle downward, atop a roughened stalk (seta), 1-1.5 cm high.
Scleropodium obtusifolium with sporophytes
Scleropodium obtusifolium sporophytes with horizontal and pendant capsules (CAH #1003) CC BY-NC Amanda Heinrich
Scleropodium obtusifolium sporophyte
Scleropodium obtusifolium sporophyte with horizontal capsule atop roughened seta (CAH #1003) CC BY-NC Amanda Heinrich
Distribution:  Scleropodium obtusifolium can be found west of the Rocky Mountains in North America. It is fairly common in seasonal creeks and streams, particularly over 2,000 feet in Santa Barbara County.
Scleropodium obtusifolium distribution map
Scleropodium obtusifolium collections. Map generated by google maps using data from the Consortium of North American Bryophyte Herbaria of accessioned collections from herbaria.
Similar Species in Santa Barbara County: Scleropodium occidentale has a nearly identical habitat preference and appearance but its leaves have a longer, stronger costa with a terminal spine.
Name: The genus name Scleropodium is derived from the Greek word skleros, meaning hard, and podion, meaning foot, and refers to the stiff seta. The species epithet, obtusifolium is a reference to the generally obtuse leaf tips.
References and Further Reading
Systematics and ecology of the moss genus Scleropodium (Brachytheciaceae), Benjamin Elias Carter, PhD Dissertation
Scleropodium obtusifolium, Michael S. Ignatov, Flora of North America Vol. 28, www.efloras.org
California Mosses, Bill and Nancy Malcolm, Jim Shevock and Dan Norris. Available for purchase from the California Native Plant Society here.
Contributions toward a Bryoflora of California II. A Key to the Mosses, Daniel H. Norris and James R. Shevock, Madrono 51(2): 133-269. 2004. Available for purchase here.
Common Mosses of Western Oregon and Washington, Bruce McCune & Martin Hutten
Plants of the Gila Wilderness